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Leaf-mine:
Rather long full depth corridor that winds freely through the leaf
and may cross itself. In the end the corridor widens considerably.
Frass mostly in a narrow central line, but may also be deposited
along the sides or be missing. The larvae regular leave a mine to
restart elsewhere. Pupation outside the mine. Neither larva nor
mine can be distinguished from that of related species (Bladmineerders
van Europa).
The
mine is also described and illustrated in British
leafminers.
Larva:
The larva is described and illustrated in British
leafminers.
Pupa:
Details unknown.
Comments:
Stachys officinalis is treated
as Betonica officinalis
(Betony) by Stace (2010).
Hosts
in Britain:
Hosts
elsewhere:
Time
of year - larvae: Spring (British
leafminers).
Time
of year - adults: Unknown.
Distribution
in Great Britain & Ireland: Widespread including Chester,
Cumberland, Derby, Dorset, Durham, East Cornwall, East Gloucester,
East Kent, East Norfolk, East Sussex, Hunts, Leicester, North Devon,
Northampton, South Northumberland, Notts, Oxford, South Hants, South-west
York, Surrey, Warwick, West Cornwall, West Gloucester and Worcester
(NBN
Gateway distribution map - BRERC,
GiGL and NE).
Also
recorded in the Republic of Ireland (Audisio in Fauna
Europaea).
NBN
Grid map:
Distribution
elsewhere: Widespread in Europe including Austria, Belgium,
Bosnia and Herzegovina, Bulgaria, Croatia, Czech Republic, Danish
mainland, French mainland, Germany, Hungary, Italian mainland, Luxembourg,
Macedonia, Poland, Portuguese mainland, Romania, Sicily, Slovakia,
Slovenia, Spanish mainland, Switzerland, The Netherlands and Ukraine
(Audisio in Fauna
Europaea).
NBN
interactive distribution maps of known host species in Britain and
elsewhere:
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Ajuga
genevensis,
Ajuga
reptans, Bellis
perennis, Clinopodium
vulgare, Digitalis
grandiflora, Digitalis
lutea, Digitalis
purpurea, Galeopsis
tetrahit, Glechoma
hederacea, Plantago
lanceolata, Plantago
major, Stachys
officinalis
(= Betonica
officinalis),
Stachys
recta, Teucrium
scorodonia, Verbascum
lychnitis, Veronica
chamaedrys
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Parasitoids
in Britain and elsewhere: Unknown.
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