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Phytomyza heracleana Hering, 1937 [Diptera: Agromyzidae] Phytomyza heracleana Hering, 1937c. Blattminen Mittel- Nordeuropas Lief 5, 6: 582. Phytomyza heracleana Hering, 1937c; Spencer, 1972b. Handbk ident. Br. Ins. 10(5g): 78 (figs 258-9), 80, 81, 121. Phytomyza heracleana Hering, 1937c; Spencer, 1976. Fauna ent. Scand. 5(1): 428, figs 747-8. Phytomyza heracleana Hering, 1937c; Spencer, 1990. Host specialization in the World Agromyzidae (Diptera) : 160, 169, 173, 175, 176 (fig. 659), 177.

Leaf-mine: A regular greenish inter-parenchymal blotch mine, frequently delimited by two veins, appearing somewhat mottled as a result of small deeper areas of feeding through the upper parenchyma (Spencer, 1972b: 78 (fig. 259); Spencer, 1976: 428, 429 (fig. 748). The mine is illustrated in Nederlandse bladmineerders. Mines of Phytomyza heracleana on Heracleum. Photos: Brian Pitkin Larva: The larva is described Griffiths (1973c), de Meijere (1937) and Ellis (Nederlandse bladmineerders). Puparium: The puparium is illustrated by Ellis (Nederlandse bladmineerders). Black; posterior spiracles each with 14-20 bulbs (Spencer, 1976: 428). Comments: Hering (1957a) includes Angelica and Seseli as host plants, but Griffiths (1973c) and Spencer (1973a) regard these records as doubtful. The species can only be distinguished from related species by characters of the male genitalia. Hosts in Britain: Ranunculaceae     Heracleum   Robbins, 1991: 67 Heracleum sphondylium Bland, 1992 Heracleum sphondylium Mines in BMNH Heracleum sphondylium Spencer, 1972b: 121 Hosts elsewhere: Ranunculaceae     ? Angelica   Spencer, 1990: 160 ? Caucalis   Spencer, 1990: 160 Heracleum   Spencer, 1990: 160 Heracleum mantegazzianum Spencer, 1976: 428 ? Heracleum mantegazzianum Nederlandse bladmineerders Heracleum sibiricum Spencer, 1976: 428 Heracleum sphondylium Spencer, 1976: 428 Heracleum sphondylium Nederlandse bladmineerders Laser   Spencer, 1990: 160 ? Laserpitium   Spencer, 1990: 160 Pastinaca   Spencer, 1990: 160 Peucedanum   Spencer, 1990: 160 Pimpinella   Spencer, 1990: 160 ? Seseli   Spencer, 1990: 160 Time of year - mines: June-August. Time of year - adults: Unknown. Distribution in Britain: Widespread including Cornwall (Portleven), Derby (Miller's Dale), Denbigh (Cefn-y-bedd), Perth (Killin) (Spencer, 1972b: 80),Inner Hebrides (Isle of Coll) (Bland, 1992), Warwick (Astley) (Robbins, 1991: 67), Linlithgow, Mid-west York, North-west York and South-west York (NBN Gateway distribution map - GiGL, LWIC and NE). Also recorded in the Republic of Ireland: Co. Clare (Burren), Co. Cork (Bantry) and Co. Kilkenny (Kilkenny) (Spencer, 1972b: 80). Distribution elsewhere: Widespread in Europe including Denmark, Norway, Sweden (Spencer, 1976: 428), The Netherlands (Nederlandse bladmineerders), Germany (Spencer, 1976: 570), Austria, Bulgaria, Czech Republic, French mainland, Hungary, Italian mainland, Lithuania and Poland (Martinez in Fauna Europaea). Parasitoids: Unknown.

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Last updated 12-Jun-2008  Brian Pitkin