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Chirosia crassiseta Stein, 1908. Wien. ent. Ztg. 27: 10.

Leaf- and stem-mine: Mine in the lowest part of the petiole, not in the pinnulae-bearing part of the rachis. Often several larvae together. Mined leaves are stunted, often incompletely rolled out. Puparium in the mine or in the ground (Bladmineerders van Europa). See de Meijere (1911a) and Brown and McGavin (1982) for details of the biology.

Larva: The larvae of flies are leg-less maggots without a head capsule (see examples). They never have thoracic or abdominal legs. They do not have chewing mouthparts, although they do have a characteristic cephalo-pharyngeal skeleton (see examples), usually visible internally through the body wall.

Larva with undivided posterior processes of the cephalopharyngeal skeleton (Bladmineerders van Europa).

Puparium: The puparia of flies are formed within the hardened last larval skin or puparium and as a result sheaths enclosing head appendages, wings and legs are not visible externally (see examples).

Hosts in Great Britain and Ireland:

Hosts elsewhere:

Dennstaedtiaceae
Pteridium	aquilinum	Bracken		Hennig, 1966-73
Pteridium	aquilinum	Bracken		Hering, 1957
Pteridium	aquilinum	Bracken		Bladmineerders van Europa

Time of year - mines: June-July (Bladmineerders van Europa).

Time of year - adults: Currently unknown.

Distribution in Great Britain and Ireland: Widespread in Britain including Berkshire, Buckinghamshire, Dorset, East Kent, Glamorgan, North Hampshire, North Somerset, North Wiltshire, Nottinghamshire, Oxfordshire, Pembrokeshire, South Devon, South-west Yorkshire, Stafford, Surrey, West Suffolk and Worcestershire (NBN Atlas).

Distribution elsewhere: Widespread in continental Europe including The Netherlands (Bladmineerders van Europa), Corsica, ? Czech Republic, Denmark, Finland, French mainland, Germany, Greek mainland, Italian mainland, Poland, Russia - Northwest, Sicily, Sweden and Yugoslavia (Michelsen in Fauna Europaea).

NBN Atlas links to known host species:

British and Irish Parasitoids in Britain and elsewhere: Currently unknown.