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Seed-feeder: Larvae feeding in seed-capsules and pupating in the seed heads (Spencer, 1990: 43).

Larva: The larvae of flies are leg-less maggots without a head capsule (see examples). They never have thoracic or abdominal legs. They do not have chewing mouthparts, although they do have a characteristic cephalo-pharyngeal skeleton (see examples), usually visible internally through the body wall.

The larva is described by Dempewolf (2001: 191).

Puparium: The puparia of flies are formed within the hardened last larval skin or puparium and as a result sheaths enclosing head appendages, wings and legs are not visible externally (see examples).

Brown, deeply segmented; posterior spiracles strongly chitinised, each with an ellipse of some 25 bulbs; all segments have a band of short spinules which are particularly strong in the anal region (Spencer, 1976: 438).

Hosts in Great Britain and Ireland:

Ranunculaceae
Aquilegia	vulgaris	Columbine		Spencer, 1972b: 118

Hosts elsewhere:

Ranunculaceae
Aquilegia				Spencer, 1990: 43
Aquilegia	vulgaris	Columbine		Spencer, 1976: 438
Aquilegia	vulgaris	Columbine		Dempewolf, 2001: 191

Time of year - larvae: Currently unknown.

Time of year - adults: Currently unknown.

Distribution in Great Britain and Ireland: Oxford (Oxford) and Monmouthshire (St. Pierre) (Spencer, 1972b: 82); East Ross, Monmouthshire and Surrey (NBN Atlas).

Distribution elsewhere: Widespread in continental Europe including Denmark, Finland, Sweden (Spencer, 1976: 438), Germany (Dempewolf, 2001: 191), Czech Republic, Norwegian mainland, Poland and The Netherlands (Fauna Europaea).

NBN Atlas links to known host species:

British and Irish Parasitoids in Britain and elsewhere: